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CATALOGUE OF NORTH AMERICAN BEETLES
OF THE
FAMILY CLERIDAE
ALBERT B. WOLCOTT
NAT.
FIELDIANA: ZOOLOGY
VOLUME 32, NUMBER 2
Published by
CHICAGO NATURAL HISTORY MUSEUM
JUNE 12, 1947
CATALOGUE OF NORTH AMERICAN BEETLES
OF THE
FAMILY CLERIDAE
ALBERT B. WOLCOTT
Assistant Curator, Harris Public School Extension
Retired February 1,
FIELDIANA: ZOOLOGY
VOLUME 32, NUMBER 2
Published by
CHICAGO NATURAL HISTORY MUSEUM
JUNE 12, 1947
THE
JUL21194*
PRINTED IN THE UNITED STATES OF AMERICA
BY CHICAGO NATURAL HISTORY MUSEUM PRESS
/ IX I
FOREWORD
My friend Albert B. Wolcott has honored me with the request
that I write a short foreword for this catalogue, and I do it with
pleasure. Wolcott, long the foremost student of the Cleridae in our
country, has established for himself a permanent place on the roster
of American Coleopterology. The present catalogue reflects his
considered opinions concerning many complicated questions of clerid
taxonomy and synonymy, with which opinions I find myself in
complete agreement. I am sure that he took no pleasure in sup-
pressing the well-known but invalidly established name, Hydnocera.
Such suppressions are, unfortunately, necessary if we are to derive
from the present confused nomenclature, based on the anarchy of
individual preference, a stable nomenclature conforming with the
International Rules.
EDWARD A. CHAPIN
United States National Museum
61
INTRODUCTION
In order that certain necessary changes in nomenclature and
systematics may be made known to those interested in the North
American Cleridae, the present revision of existing catalogues is
offered.
It was not until 1841 that an attempt was made to bring together
the available knowledge concerning the group of beetles now known
as the family Cleridae. In that year the Marquis Spinola published
a synopsis of the genera that he considered as belonging to the group,
with indications that a monographic treatment of the species would
follow. Apparently stimulated by this preliminary paper, Klug
(1842) gave the descriptions of the new species and genera known
to him. In 1844, there appeared the promised Spinola monograph,
a beautifully prepared and fully illustrated account of the world
fauna of this interesting family. Thus a foundation was laid for the
further study of the Cleridae.
Five years later LeConte (1849) offered the first synopsis of the
North American species, including therein many not previously
known. Further elucidation of our fauna was later made by the
same author (1861) and by LeConte and Horn (1883); both papers
were published by the Smithsonian Institution.
During this period, various synopses and catalogues of the world
fauna had appeared. Most notable among these is the generic
synopsis of Lacordaire (Gen. Coleopt., 4, 1857). In 1849 Adam
White prepared and issued a catalogue of the coleopterous insects
in the British Museum, describing the new genera and species found
in that collection. Two papers of similar scope were written by
Chevrolat (1874, 1876) dealing with his own collection. In 1869
there appeared the first world catalogue of the Coleoptera, by
Gemminger and Harold. At this date 697 species of clerids were
known from the whole world.
Henry S. Gorham was the author of a series of papers on the
family, starting in 1876 and continuing into the 1900's, among
which is the part on the Cleridae in the Biologia Centr all- Americana
(1882-86). This contribution is of definite interest to North Ameri-
can students.
63
64 FIELDIANA: ZOOLOGY, VOLUME 32
In 1900, Lodhe's catalogue appeared (Stettin, ent. Zeitg., 61,
1900). It lists a total of 1,822 species, a testimonial to the industry
of the species describers during the previous thirty years. The need
for a new generic revision was obvious and this need was met by the
appearance in 1903 of Schenkling's masterful work in the Wytsman
Genera Insectorum. Such a work always stimulates activity in a
group, and Schenkling (1906) found it necessary to introduce into
the literature many changes in systematics and nomenclature, in
anticipation of his forthcoming part of the Junk Catalogue Coleop-
terorum. This was published in 1910 and showed that the number
of species of clerids had increased to 2,285. Since the publication
of the 1910 catalogue, a paper of general interest, dealing with the
systematics of the group as a whole, has been published by Chapin
(1924). In it the subfamilies are discussed and re-defined. Students
of the American fauna will also be interested in a manual of the
American genera of Coleoptera prepared by Bradley (1930).
In recent years some important subfamily and generic revisions
have appeared. The more important of these are Linsley's study
of the genus Aulicus (1936), Corporaal's revision of the subfamily
Thaneroclerinae (1939), and Wolcott's treatment of the genus
Orthopleura (1943).
It is necessary to explain my reason for considering Clerus
alvearius Fabricius to be the genotype of Trichodes Herbst. Malaise
(Ent. News, 48, pp. 130-134, 1937; Ent. Tidskr., 59, pp. 99-106,
1938) and Blackwelder (Bull. Brooklyn Ent. Soc., 41, pp. 72-78,
1946) have outlined the arguments for accepting Fabricius as the
designator of genotypes; McAtee and Malloch (Proc. U. S. Nat.
Mus., 67, Art. 1 [No. 2573], pp. 129-131, 1926) have given argu-
ments against this point of view. I am in agreement with Malaise
and Blackwelder and consider that Clerus alvearius was established
as the genotype of Trichodes by Fabricius (Syst. Eleuth., 1, p. 284,
1801). If, in the future, the Commission renders a definite opinion
against accepting the Fabrician designations, then the name Tri-
chodes must be suppressed and the name Clerus applied to the genus,
since Curtis (Brit. Ent., 1, pi. 44, 1824) later established alvearius
as the genotype of Clerus.
In closing, I may say that no apologies are offered for the elimina-
tion of the well-known generic name Hydnocera from the family.
The change is dictated by the Rules of the International Commission
on Zoological Nomenclature. Dejean, in his series of catalogues,
proposed numerous new generic names, many of which are defined
WOLCOTT: FAMILY CLERIDAE 65
by the inclusion of species which had already been validly described.
Students of the Rules agree that such names fulfill the requirements
of the Code and must be accepted as valid. However, for many
years the Dejean catalogues were treated by other writers as a
source of names, and many cases of transference of names have
occurred. Phyllobaenus was proposed by Dejean to include, among
others, Clerus humeralis Germar, a validly described species. Spinola
disregarded this and transferred the name to a genus in another
subfamily of the Cleridae. Under the Rules it must be restored to
its rightful position, making Hydnocera Newman a synonym.
I offer thanks to Dr. Edward A. Chapin, Curator of Insects,
United States National Museum, for reading this manuscript and
making certain suggestions.
ALBERT B. WOLCOTT
FAMILY CLERIDAE
Subfamily TILLINAE
Genus Monophylla Spinola
Monophylla Spinola, Rev. Zool., 4, p. 75, 1841; Monog. C16r., 1, p. 384,
1844.
Macrotelus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, p. 274, 1842.
Elasmocerus LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 13, 1849.
Genotype: Tillus terminatus Say
californica Fall, Occ. Papers Calif. Acad. Sci., 8, p. 251, 1901. Utah, Oregon,
substriata Wolcott, Field Mus. Nat. Hist., Zool. Ser.,
7, p. 340, 1910.
pallipes Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 128, 1908.
ruficollis Schaeffer, op. cit., 19, p. 121, 1911.
Costa Rica
Texas
Arizona
terminata Say, Boston Journ. Nat. Hist., 1, p. 160, 1835. Eastern North
megatoma Spinola, Monog. Cler., 1, p. 385, pi. 28, fig. 5,
1844'
Arizona
Genus Callotillus- Wolcott
Callotillus Wolcott, Ent. News, 22, p. 115, 1911.
Genotype: Callotillus eburneocinctus Wolcott
eburneocinctus Wolcott, loc. cit., p. 115.
elegans Erichson, Arch. Naturg., 13, p. 85, 1847.
occidentalis Gorham, Biol. Centr.-Amer., Ins.-Coleop.,
3, pt. 2, p. 129, pi. 9, figs. 1, 2, 1882.
vafer Wolcott, Proc. U. S. Nat. Mus., 59, p. 270, 1921.
Southern Flor-
ida
Texas, New
Central Amer-
ica
California, Ari-
zona, New
Mexico
Genus Tillus Olivier
Tillus Olivier, Ent., 2, No. 22, 1790.
Cylinder Voet, Cat. Syst. Col., 1, p. 78, 1769-1806 (not Mueller 1766).
Genotype: Chrysomela elongata Linnaeus, Syst. Nat., ed. 10, p. 377, 1758
(Europe)
67
68
FIELDIANA: ZOOLOGY, VOLUME 32
collaris Spinola, Monog. Cle"r., 1, p. 98, pi. 2, fig. 6, 1844. Ohio, Georgia,
Louisiana,
Alabama,
Texas, Mexico
patagoniae Knull, Ohio Journ. Sci., 46, p. 72, 1946.
Arizona
Genus Perilypus Spinola
Perilypus Spinola, Monog. Cler., 1, p. 103, 1844.
Genotype: Perilypus carbonarius Spinola
carbonarius Spinola, loc. cit., p. 105, pi. 5, fig. 4.
California,
Nevada
Genus Lecontella Wolcott and Chapin
Lecontella Wolcott and Chapin, Bull. Brooklyn Ent. Soc., 13, p. 107, 1918.
Genotype: Cymatodera cancellata Leconte
cancellata LeConte, Proc. Acad. Nat. Sci. Phila., 7, p. 81,
1854.
brunnea Melsheimer, Proc. Acad. Nat. Sci. Phila., 2,
p. 306, 1846 (not Spinola 1844).
gnara Wolcott, Coleop. Contr., 1, p. 105, 1927.
Maine, North
Carolina, Mis-
Texas, Arizona
Oklahoma,
Arizona
Genus Cymatodera Gray
Cymatodera Gray, in Griffith's Anim. Kingd., Ins., 1, p. 375, 1832.
Cymatoderus Desmarest in Chenu, Encycl. d'Hist. Nat., Coleop., 2, p. 240,
1860.
Genotype: Cymatodera hopei Gray
aegra Wolcott, Proc. U. S. Nat. Mus., 59, p. 271, pi. 43, Arizona
figs. 2, 3, 1921.
aemula Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 348, Arizona
pi. 6, fig. 9, 1910.
Texas, Arizona
California,
Mexico,
Arizona
aethiops Wolcott, loc. cit., p. 350, pi. 6, figs. 10-13.
angustata Spinola, Monog. Cle>., 1, p. 149, pi. 7, fig. 1, 1844.
langustata LeConte, Smiths. Misc. Coll., 6, No. 167,
2d ed., p. 96, 1866.
antennata Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 128, 1908. Arizona
balteata LeConte, Proc. Acad. Nat. Sci. Phila., 7, p. 81, 1854. Texas
fallax Horn, Trans. Amer. Ent. Soc., 12, p. 153, 1885.
belfragei Horn, op. cit., 5, p. 226, pi. 1, fig. 11, 1876. Texas
bicolor Say, Journ. Acad. Nat. Sci. Phila., 5, p. 174, 1825. Canada,
Georgia, New
York, Illinois,
Kansas, Texas,
Arizona
WOLCOTT: FAMILY CLERIDAE
brevicollis Schaeffer, Journ. N. Y. Ent. Soc., 25, p. 130, 1917.
californica Horn, Trans. Amer. Ent. Soc., 2, p. 134, 1868.
cephalica Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 130, 1908.
cognata Wolcott, Ent. News, 21, p. 320, 1910.
confusa Wolcott, Proc. U. S. Nat. Mus., 59, p. 282, 1921.
cylindricollis Chevrolat, Coleop. Mex., Cent. I, Fasc. 1, No.
11, 1833.
decipiens Fall, Can. Ent., 38, p. 114, 1906.
delicatula Fall, loc. c^t., p. 113.
fascifera LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed.,
p. 95, 1866.
flavosignata Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 129,
1908.
fuchsii Schaeffer, op. cit., 12, p. 216, 1904.
comans Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7,
p. 351, pi. 6, figs. 14-16, 1910.
fuscula LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 212, 1852.
var. texana Gorham, Biol. Centr.-Amer., Ins.-Coleop., 3,
pt. 2, p. 134, 1882.
hopei Gray in Griffith's Anim. Kingd., Ins., 1, p. 375, pi. 48,
fig. 1, 1832.
gigantea Horn, Trans. Amer. Ent. Soc., 8, p. 148, 1880.
horni Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 349,
1910; Journ. N. Y. Ent. Soc., 30, pp. 191-194, figs.
1, 2, 1922.
californica Horn, Trans. Amer. Ent. Soc., 5, p. 225, pi. 1,
fig. 9, 1876 (not Horn 1868).
inornata Say, Boston Journ. Nat. Hist., 1, p. 161, 1835.
tenera LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 14,
1849.
isabellae Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 345,
pi. 6, fig. 7, 1910.
knausi Wolcott, Proc. U. S. Nat. Mus., 59, p. 278, pi. 43,
figs. 24, 25, 1921.
laevicollis Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 130, 1908.
latefascia Schaeffer, op. cit.; 12, p. 216, 1904.
longicornis LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 16,
1849.
Arizona
California,
Lower Cali-
fornia
Lower Cali-
fornia
Utah, Nevada
North Carolina,
Missouri,
Texas
Arizona, Mex-
ico
California
Arizona, Cali-
fornia, Lower
California,
Mexico
Lower Cali-
fornia
Arizona
Utah, Nevada,
Texas, Arizona,
New Mexico,
California
Arizona, Mex-
ico
Texas
California,
Mexico
Arizona, Cali-
fornia
Canada, East-
ern United
States to Utah
and Arizona
Utah, Arizona
Utah, Arizona
Arizona
Arizona
Utah, Arizona,
New Mexico
70 FIELDIANA: ZOOLOGY, VOLUME 32
longula Wolcott, Proc. U. S. Nat. Mus., 59, p. 280, 1921. Arizona
rudis Wolcott, loc. cit., p. 281.
mitchelli Chapin, Proc. Biol. Soc. Wash., 40, p. 144, 1927. Texas, Mexico
mitis Wolcott, Proc. U. S. Nat. Mus., 59, p. 273, pi. 43, fig. 8, Arizona
1921.
morosa LeConte, Proc. Acad. Nat. Sci. Phila., 10, p. 71, 1858. Colorado, Ari-
zona, New
Mexico, Cali-
fornia, Mexico
mystica Wolcott, Proc. U. S. Nat. Mus., 59, p. 275, pi. 43, Utah
figs. 16-19, 1921.
neomexicana Knull, Ent. News, 45, p. 9, 1934. New Mexico
obliquefasciata Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 215, New Mexico,
1904. • Texas
oblita Horn, Trans. Amer. Ent. Soc., 5, p. 227, pi. 1,' fig. 12, Nevada, Ari-
1876. zona, Lower
California
ovipennis LeConte, Proc. Acad. Nat. Sci. Phila., 11, p. 76, Arizona, New
1859. Mexico, Cali-
pilosella LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed., forma
p. 95, 1866.
pallida Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 128, 1908. Texas
peninsularis Schaeffer, op. cit., 12, p. 214, 1904. Arizona, Lower
California
pubescens Wolcott, Bull. Wise. Nat. Hist. Soc., 7, p. 94, 1909. Texas
punctata LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 212, California,
1852. Lower Cali-
fornia
puncticollis Bland, Proc. Ent. Soc. Phila., 1, p. 356, 1863. Texas, New
Mexico, Lower
California
purpuricollis Horn, Proc. Calif. Acad. Sci., 4, p. 381, 1894. Arizona, Lower
California
rufiventris Wolcott, Coleop. Contr., 1, p. 105, 1927. California
santarosae Schaeffer, Bull. Brooklyn Mus., 1, No. 7, p. 152, Lower Cali-
1905. fornia
schwarzi Wolcott, Proc. U. S. Nat. Mus., 59, p. 274, pi. 43, Arizona
figs. 11, 12, 1921.
sirpata Horn, Trans. Amer. Ent. Soc., 12, p. 152, 1885. Texas
var. spatiosa Wolcott, Proc. U. S. Nat. Mus., 59, p. 277, Texas
pi. 43, figs. 22, 23, 1921.
snow! Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 349, Arizona
1910.
soror Wolcott, loc. cit., p. 342, pi. 6, figs. 1, 2. Arizona
subsimilis Wolcott, loc. cit., p. 343, pi. 6, figs. 3, 4. Arizona
WOLCOTT: FAMILY CLERIDAE
torosa Wolcott, loc. cit., p. 347, pi. 6, fig. 8.
tricolor Skinner, Ent. News, 16, p. 292, 1905.
turbata Horn, Trans. Amer. Ent. Soc., 12, p. 151, 1885.
tuta Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 344, pi. 6,
figs. 5, 6, 1910. .
umbrina Fall, Can. Ent., 38, p. 114, 1906.
undulata Say, Journ. Acad. Nat. Sci. Phila., 5, p. 174, 1825.
bosci Chevrolat, Ann. Soc. Ent. France, (2), 1, p. 31, 1843.
longicollis Spinola, Monog. Cler., 1, p. 147, pi. 10, fig. 1,
1844.
var. arizonica Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 130,
1908.
var. brunnea Spinola, Monog. Cle>., 1, p. 148, 1844.
var. cauthieri Spinola, loc. cit., p. 147.
uniformis Schaeffer, Bull. Brooklyn Mus., 1, No. 7, p. 151,
1905.
usta LeConte, Proc. Acad. Nat. Sci. Phila., 10, p. 71, 1858.
vandykei Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 217, 1904.
xanti Horn, Trans. Amer. Ent. Soc., 5, p. 222, pi. 1, fig. 3, 1876.
xavierae Knull, Ann. Ent. Soc. Amer., 33, p. 555, fig. 1, 1940.
71
Arizona, New
Mexico
Arizona
Texas, Arizona
Arizona, New
Mexico
Colorado, New
Mexico
Canada, Mary-
land, Illinois,
Kentucky,
Arkansas,
Texas, Arizona
Arizona
Arizona
"North Amer-
ica"
Arizona
Texas, Arizona,
New Mexico
California
Lower Cali-
fornia, .Mexico
Arizona
Genus Bostrichoclerus Van Dyke
Bostrichoclerus Van Dyke, Ent. News, 49, p. 189, 1938.
Genotype : Bostrichoclerus bicornus Van Dyke
bicornus Van Dyke, loc. cit., p. 190.
Gulf of Cali-
fornia
Subfamily PHYLLOBAENINAE
Genus Phyllobaenus Dejean
Phyllobaenus Dejean, Cat. Coleop., ed. 3 (rev.), p. 127, 1837.
Hydnocera Newman, Ent. Mag., 5, p. 379, 1838.
Theano Chevrolat, Ann. Soc. Ent. France, (2), 1, p. 33, 1843 (not Castelnau
1836).
Genotype: Clerus humeralis Say
affiliatus Fall, Can. Ent., 38, p. 117, 1906. California
arizonicus Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 132, 1908. Arizona
bicolor LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 213, 1852. California
72
FIELDIANA: ZOOLOGY, VOLUME 32
bimaculatus Wolcott, Can. Ent., 40, p. 232, 1908.
binotatus Chapin, Proc. Biol. Soc. Wash., 31, p. 107, 1918.
blanchardi Wolcott, Ent. News, 39, p. 210, 1928.
chapini Wolcott, Coleop. Contr., 1, p. 81 (footnote), 1927.
lateralis Schaeffer, Bull. Brooklyn Mus., 1, p. 157, 1905
(not Gorham 1883).
cinctus Spinola, Monog. Cle>., 2, p. 48, pi. 39, fig. 5, 1844.
cobaltinus Chapin, Proc. Biol. Sac. Wash., 35, p. 55, 1922
(as colbaltina).
coeruleipennis Wolcott, Can. Ent., 40, p. 229, 1908.
commixtus Chapin, Proc. Biol. Soc. Wash., 35, p. 58, 1922.
cribripennis Fall, Can. Ent., 38, p. 116, 1906.
cuneiformis Wolcott, Ent. News, 39, p. 208, 1928.
cyanitinctus Fall, Can. Ent., 38, p. 116, 1906.
discoideus LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 213,
1852.
furcata Gorham, Biol. Centr.-Amer., Ins.-Coleop., 3,
pt. 2, p. 342, pi. 13, fig. 14, 1886.
dubius Wolcott, Bull. Lab. Nat. Hist. Univ. Iowa, 6, p. 60,
1912.
fallax Wolcott, Can. Ent., 40, p. 230, 1908.
fuchsi Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 132, 1908.
nunnenmacheri Schaeffer, loc. cit., p. 133.
gahani Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 379,
1910.
hamatus LeConte, Ann. Kept. Chief Engrs. App. JJ, p. 517,
1876.
humeralis Say, Journ. Acad. Nat. Sci. Phila., 3, p. 192, 1823.
var. cyanescens LeConte, Ann. Lye. Nat. Hist. N. Y., 5,
p. 28, 1849.
difficilis LeConte, loc. cit., p. 27.
iowensis Chapin, Proc. Biol. Soc. Wash., 35, p. 57, 1922.
knausi Wickham, Can. Ent., 37, p. 168, 1905.
lecontei Wolcott, Bull. Lab. Nat. Hist. Univ. Iowa, 6, p. 62,
1912.
subaenea LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 26,
1849 (not Spinola 1844).
longus LeConte, Trans. Amer. Ent. Soc., 12, p. 22, 1884.
California
California
New Mexico
Texas
Arizona, Cen-
tral and South
America
California
Arizona
Massachusetts,
New York
Texas
Arizona
New Mexico
Arizona, New
Mexico, Utah,
California,
Mexico
North Dakota
Colorado
Arizona, New
Mexico
Texas
New Mexico
Maine to Flor-
ida, Illinois,
Kansas
Texas, Arizona
Iowa
Kansas, Texas
Maine to
Louisiana,
Arizona, Alaska
Arizona
WOLCOTT: FAMILY CLERIDAE
73
maritimus Wolcott, Ent. News, 21, p. 321, 1910.
mirus Wolcott, op. cit., 39, p. 207, 1928.
nigrescens Schaeffer, Bull. Brooklyn Mus., 1, No. 15, p. 381,
1909.
niveifasciatus Schaeffer, op. cit., 1, No. 7, p. 156, 1905.
occidentalis Chapin, Proc. Biol. Soc. Wash., 35, p. 56, 1922.
omoger Horn, Proc. Calif. Acad. Sci., 4, p. 383, 1894.
pallipennis Say, Journ. Acad. Nat. Sci. Phila., 5, p. 176, 1825.
serrata Newman, Ent. Mag., 5, p. 379, 1838.
picipennis Chapin, Proc. Biol. Soc. Wash., 35, p. 56, 1922.
plagifer Fall, Can. Ent., 38, p. 115, 1906.
pubescens LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 28, 1849.
pulcher Chapin, Proc. Biol. Soc. Wash., 35, p. 57, 1922.
puritanus Wolcott, Ent. News, 39, p. 208, 1928.
pygmaeus Wolcott, Bull. Lab. Nat. Hist. Univ. Iowa, 6, p. 63,
1912.
quadrimaculatus Van Dyke, Bull. Brooklyn Ent. Soc., 10,
p. 31, 1915.
robustus Horn, Trans. Amer. Ent. Soc., 2, p. 135, 1868.
rufipes Newman, Mag. Nat. Hist., (2), 4, p. 363, 1840.
scaber LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 213, 1852.
funebris Chevrolat, Rev. Mag. Zool., (3), 2, p. 321, 1874.
simulans Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 133, 1908.
singularis Wolcott, Bull. Lab. Nat. Hist. Univ. Iowa, 6, p. 62,
1912.
subaeneus Spinola, Monog. Cle>., 2, p. 51, 1844.
steniformis Spinola, loc. cit., p. 144, pi. 40, fig. 2 (not
steniformis Klug 1842).
subfasciatus LeConte, Smiths. Misc. Coll., 6, No. 167, 2d
ed., p. 97, 1866.
var. fraterna Wolcott, Bull. Lab. Nat. Hist. Univ. Iowa, 6,
p. 61, 1912.
suturalis Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42,
p. 313, 1842.
Massachusetts
Nebraska
North Carolina
Arizona,
Mexico
California
Arizona, Cali-
fornia, Lower
California
Canada, Ala-
bama, Colorado
Texas
California
Itlinpis, Mis-
souri, Kansas,
Oklahoma,
Colorado, Texas
California
Massachusetts
California
California
California
Florida, Ala-
bama, Arkansas
Kansas, Ari-
zona, New
Mexico,
Wyoming,
Idaho,
Washington
Arizona
North Carolina
Massachusetts
Kansas,
Nebraska,
Utah, Texas,
California
Massachusetts
Florida
u. OF iu- ua
74
FIELDIANA: ZOOLOGY, VOLUME 32
limbata Spinola, Monog. Cler., 2, p. 49, 1844.
Imarginata var. Gorham, Biol. Centr.-Amer., Ins.-
Coleop., 3, pt. 2, p. 168, 1882.
var. aequinoctialis Spinola, Monog. Cler., 2, p. 50, 1844.
tibialis Wolcott, Ent. News, 21, p. 322, 1910.
tricolor Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 219, 1904.
tristis Schaeffer, Bull. Brooklyn Mus., 1, No. 15, p. 381, 1909.
unifasciatus Say, Journ. Acad. Nat. Sci. Phila., 5, p. 176,
1825.
punctate, Spinola, Monog. Cle>., 2, p. 54, pi. 40, fig. 4,
1844.
verticalis Say, Boston Journ. Nat. Hist., 1, p. 164, 1835.
brachyptera Klug, Abh. Konig. Akad. Wiss. Berlin,
1840-42, p. 313, 1842.
lineatocollis Spinola, Monog. Cler., 2, p. 51, pi. 40, fig. 1,
1844.
tenella Say, Boston Journ. Nat. Hist., 1, p. 164, 1835.
vicinus Chapin, Proc. Biol. Soc. Wash., 35, p. 55, 1922.
wickhami Wolcott, Can. Ent., 40, p. 231, 1908.
Mexico
Mexico
Nebraska
Texas
Arizona,
Mexico
Canada, Maine,
Illinois,
Alabama,
Kansas, Colo-
rado, Okla-
homa, Arizona
Canada, New
Hampshire,
Illinois, Florida,
Minnesota,
South Dakota,
Kansas, Texas
Texas
Arizona
Genus Isohydnocera Chapin
Isohydnocera Chapin, Bull. Brooklyn Ent. Soc., 12, p. 83, 1917.
Genotype: Hydnocera curtipennis Newman
aegra Newman, Mag. Nat. Hist., (2), 4, p. 364, 1840.
albocincta Horn, Trans. Amer. Ent. Soc., 3, p. 342, 1871.
brunnea Chapin, Bull. Brooklyn Ent. Soc., 12, p. 84, 1917.
curtipennis Newman, Mag. Nat. Hist., (2), 4, p. 364, 1840.
longicollis Ziegler, Proc. Acad. Nat. Sci. Phila., 2, p. 44,
1844.
gerhardi Wolcott, Ent. News, 21, p. 323, 1910.
liebecki Wolcott, op. cit., 39, p. 211, 1928.
mima Wolcott, loc. cit.
nigrina Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 134, 1908.
ornata Wolcott, Can. Ent., 40, p. 233, 1908.
pusilla Schaeffer, Bull. Brooklyn Mus., 1, No. 15, p. 381, 1909.
schusteri LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed.,
p. 97, 1866.
Georgia,
Florida
Texas
Kansas, Texas
Canada, Con-
necticut, Dis-
trict of Colum-
bia, Illinois,
Indiana,
Minnesota
Arizona
New Jersey
New Mexico
Arizona
Arizona
Arizona
Illinois
WOLCOTT: FAMILY CLERIDAE
tabida LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 29, 1849.
tricondylae LeConte, loc. cit., p. 26.
Genus Wolcottia Chapin
Wolcotlia Chapin, Bull. Brooklyn Ent. Soc., 12,' p. 84, 1917.
Genotype: Hydnocera pedalis LeConte
parviceps Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 134, 1908.
pedalis LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed., p. 97,
1866.
sobrina Fall, Can. Ent., 38, p. 117, 1906.
75
Ontario, Wis-
consin, Illinois,
Indiana, Ala-
bama, Kansas
Illinois, Mis-
souri, Colorado,
Nebraska,
Arizona
Arizona
New Jersey,
Illinois,
Wisconsin,
Missouri,
Nebraska,
Arizona
Arizona
Subfamily THANEROCLERINAE1
Genus Thaneroclerus Lefebvre
Thaneroclerus Lefebvre, Bull. Soc. Ent. France, (1), 7, p. 13, 1838.
Thaneclerus Desmarest in Chenu, Encycl. d'Hist. Nat., Coleop., 2, p. 247, 1860.
T hanater oclerus Gemminger and Harold, Cat. Coleop., 6, p. 1739, 1869.
Taneroclerus Chevrolat, Bull. Soc. Ent. France, (5), 10, p. 31, 1880.
Metademius Matsumura, Dainippon Gaichuzensho, 2, p. 234, (?)1935 (not
Schenkling 1899).
Genotype: Clerus buquet Lefebvre
buquet Lefebvre, Ann. Soc. Ent. France, 4, p. 577, pi. 16,
fig. 4, 1835.
buqueti Westwood, Bull. Soc. Ent. France, (1), 7, p. 13,
1838.
girodi Chevrolat, op. cit., (5), 10, p. 31, 1880.
tabacci Matsumura, Dainippon Gaichuzensho, 2, p. 234,
(?)1935.
var. dermestoides Klug, Abh. Konig. Akad. Wiss. Berlin,
1840-42, p. 310, 1842.
Florida, New
York, Califor-
nia, cosmopoli-
tan
New York
Genus Zenodosus Wolcott
Zenodosus Wolcott, Ent. News, 21, p. 321, 1910.
1 A revision of this subfamily is given by J. B. Corporaal (1939).
76
FIELDIANA: ZOOLOGY, VOLUME 32
Xenodosus Champlain, Proc. U. S. Nat. Mus., 57, p. 636, 1920.
Zenodusus Leng, Cat. Coleop. N. Amer., p. 470, 1920 (err. typ.).
Genotype: Clerus sanguineus Say
sanguineus Say, Boston Journ. Nat. Hist., 1, p. 162, 1835.
Ontario, Maine,
south to North
Carolina and
Arkansas, west
to Colorado
Genus Ababa Casey
Ababa Casey, Ann. N. Y. Acad. Sci., 9, p. 653, 1897.
Prionodera Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 396, 1910.
Prionostichaeus Wolcott, Ent. News, 22, p. 125, 1911.
Wolcottella Lucas, Arch. Naturg., 84, Abt. A, p. 669, 1918 (1920).
Genotype: Clerus tantillus LeConte, through synonymy
tantilla LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed., p. 96,
1866.
crinita Casey, Ann. N. Y. Acad. Sci., 9, p. 654, 1897.
Massachusetts,
District of
Columbia, Mis-
souri, Alabama,
Louisiana^
Texas
Subfamily CLERINAE
Genus Priocera Kirby
Priocera Kirby, Trans. Linn. Soc. London, 12, p. 389, 1818.
Genotype: Priocera variegata Kirby, Trans. Linn. Soc. London, 12, p. 392,
1818 (Brazil)
castanea Newman, Ent. Mag., 5, p. 380, 1838. Connecticut,
maculata Ziegler, Proc. Acad. Nat. Sci. Phila., 2, p. 268, ES
iQjr nois,
Michigan,
rufescens Spinola, Monog. Cler., 1, p. 119, pi. 4, fig. 3, Kansas
1844.
catalina Cazier, Pan-Pac. Ent., 15, p. 162, 1939.
Arizona
chiricahuae Knull, Ann. Ent. Soc. Amer., 32, p. 27, pi. 1, Arizona
fig. 3, 1939.
lecontei Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 356, California
pi. 5, fig. 2, 1910.
pusilla Kirby, Zool. Journ., 2, p. 518, 1826. "North Amer-
ica"
Genus Colyphus Spinola
Colyphus Spinola, Rev. Zool., 4, p. 72, 1841; Monog. Cler., 1, p. 133, 1844.
Ckronomus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, p. 282, 1842.
WOLCOTT: FAMILY CLERIDAE 77
Derestenus Chevrolat, Guer. Mag. Zool., (2), 5, p. 13, 1843.
Genotype: Colyphus signaticollis Spinola, Monog. Cle>., 1, p. 135, pi. 5, fig. 5,
1844 (Central America)
furcatus Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 218, 1904. Texas
ornaticollis LeConte, Trans. Amer. Ent. Soc., 8, p. 194, 1880. Ohio
melanopterus Dury, Journ. Cincinn. Soc. Nat. Hist., 20,
p. 251, 1906.
Genus Placopterus Wolcott
Placopterus Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 363, 1910.
Poecilochroa Chevrolat, Mem. Cler., p. 5, 1876 (not Westring 1874).
Ploeopterus Leng, Cat. Coleop. N. Amer., p. 149, 1920 (err. typ.).
Phloeopterus Britton, Bull. Conn. Geol. Nat. Hist. Surv., 31, p. 236, 1931
(err. typ.).
Genotype: Poecilochroa plumbea Gorham, Biol. Centr.-Amer., Ins.-Coleop., 3,
pt. 2, p. 337, pi. 13, fig. 11 (Mexico)
cyanipennis Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, Mexico
p. 307, 1842.
var. dasytoides Westwood in White, Nomencl. Coleop. Ins. Mexico, Texas
Brit. Mus., 4, p. 50, 1849.
haagi Chevrolat, Mem. Cler., p. 12, 1876. Texas, Mexico
subcostatus Schaeffer, Journ. N. Y. Ent. Soc., 25, p. 131, Florida, Texas
1917.
thoracicus Olivier, Ent., 4, No. 76, p. 18, pi. 2, fig. 22, 1795. Canada to
monilis Melsheimer, Proc. Acad. Nat. Sci. Phila., 2, S^feSh
r» 9fi7 1 9.AK. HOIS, inUldCd,
p. 307, 1846. California,
Texas, Mexico
var. pallipes Wolcott, Bull. Lab. Nat. Hist. Univ. Iowa, 6,
p. 55, 1912.
Genus Thanasimus Latreille
Thanasimus Latreille, Gen. Crust, et Ins., 1, p. 270, 1806.
Cleroides Schaeffer, Elem. Ent., pi. 137, 1777.
Pseudoclerus Jacq. du Val, Gen. Col. d'Eur., 3, p. 196, 1860.
Genotype: Attelabus formicarius Linnaeus, Syst. Nat., ed. 10, p. 387, 1758
(Europe)
dubius Fabricius, Gen. Ins., p. 229, 1777. Maine to
ruficeps Spinola, Monog. Cle>., 1, p. 189, pi. 14, fig. 3,
Louisiana,
Kansas,
Nebraska,
Colorado, New
Mexico '
78
FIELDIANA: ZOOLOGY, VOLUME 32
repandus Horn, Trans. Amer. Ent. Soc., 3, p. 342, 1871.
trifasciatus Say, Journ. Acad. Nat. Sci. Phila., 5, p. 175, 1825.
nigripes var. White, Nomencl. Coleop. Ins. Brit. Mus., 4,
p. 23, 1849.
undatulus Say, Boston Journ. Nat. Hist., 1, p. 163, 1835.
undulatus Leng. et auct. (err. typ.).
var. abdominalis Kirby, Fauna Bor.-Amer., 4, p. 244, pi. 2,
fig. 5, 1837.
var. nubilus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42,
p. 386, 1842.
var. melanocephalus Chevrolat, Rev. Mag. Zool., (3), 2,
p. 290, 1874.
var. monticola Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7,
p. 357, pi. 5, fig. 3, 1910.
var. rubriventris LeConte, Proc. Acad. Nat. Sci. Phila., 13,
p. 351, 1861M
California
Canada, Ver-
mont, Wiscon-
sin, Pennsyl-
vania
Canada, Maine,
west to Mon-
tana and south
to Tennessee
and New
Mexico
Nova Scotia,
New Hamp-
shire
British Colum-
bia, Washing-
ton, California
California
Genus Enoclerus Gahan
Enoclerus Gahan, Ann. Mag. Nat. Hist., (8), 5, pp. 62, 65, 1910.
Clerus auctorum (not Fabricius 1775).
Genotype: Clerus sexguttatus Fabricius, Syst. Ent., p. 823, 1775 (South
America)
abdominalis Chevrolat, Coleop. Mexico, Cent. I, Fasc. 3,
No. 52, 1834.
zonatus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42,
p. 297, 1842.
var. spinolae LeConte, Proc. Acad. Nat. Sci. Phila., 6,
p. 230, 1853.
acerbus Wolcott, Ent. News, 22, p. 119, 1911.
analis LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 20, 1849.
badeni Gorham, Cist. Ent., 2, p. 80, 1876.
bimaculatus Skinner, Ent. News, 16, p. 291, 1905.
bioculatus Schaeffer, Bull. Brooklyn Mus., 1, No. 7, p. 154,
1905 (err. typ.).
Mexico
Kansas, Utah,
California,
Nevada, New
Mexico, Ari-
zona, Texas,
Mexico
Utah, Nevada
Ohio, Illinois,
Kansas, Colo-
rado, Califor-
nia, New
Mexico, Ari-
zona, Texas,
Mexico
Texas, Cali-
fornia, Mexico
Arizona
WOLCOTT: FAMILY CLERIDAE
79
bombycinus Chevrolat, Coleop. Mexico, Cent. I, Fasc. 1,
No. 12, 1833.
aeneicollis Spinola, Monog. Cle>., 1, p. 254, pi. 24, fig. 1,
1844.
sericans Westwood in White, Nomencl. Coleop. Ins.
Brit. Mus., 4, p. 58,' 1849.
brevicollis Spinola, Monog. Cler., 1, p. 266, pi. 25, fig. 5, 1844.
chapini Wolcott, Trans. Amer'. Ent. Soc., 48, p. 74, 1922.
coccineus Schenkling, Deutsch. Ent. Zeitschr., 1906, p. 272,
pi. 2, fig. 7, 1906.
corallinus Fall, Trans. Amer. Ent. Soc., 33, p. 240, 1907.
cordifer LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 21, 1849.
crabronarius Spinola, Monog. Cler., 1, p. 247, pi. 23, fig. 1,
1844.
humeralis Schaeffer, Bull. Brooklyn Mus., 1, No. 7, p. 155,
1905.
ichneumoneus Fabricius, Gen. Ins., p. 230, 1777.
rufus Olivier, Encycl. Meth., 6, p. 13, 1791.
var. knabi Wolcott, Ent. News, 21, p. 321, 1910.
inyoensis Van Dyke, Ent. News, 49, p. 191, 1938.
laetus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, p. 301,
1842.
abruptus LeConte, Proc. Acad. Nat. Sci. Phila., 10, p. 72,
1858.
lautus Wolcott, Trans. Amer. Ent. Soc., 48, p. 69, 1922.
lecontei Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 359,
1910.
nigriventris LeConte, Proc. Acad. Nat. Sci. Phila., 13,
p. 351, 1861 (not nigriventris Blanchard 1843).
California,
Mexico.
North America
Arizona
Minnesota to
Texas, Mexico
Montana,
Kansas, Colo-
rado, Nebraska,
New Mexico,
Texas
? Texas
cupressi Van Dyke, Bull. Brooklyn Ent. Soc., 10, p. 29, 1915. California
decussatus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, Arizona,
p. 296, 1842. Mexico
hopfneri Spinola, Monog. Cler., 1, p. 256, 1844.
erro Wolcott, Trans. Amer. Ent. Soc., 48, p. 68, 1922. Arizona
eximius Mannerheim, Bull. Soc. Imp. Nat. Moscou, 16, p. 248, California
1843.
holosericeus White, Nomencl. Coleop. Ins. Brit. Mus., 4,
p. 58, 1849.
gerhardi Wolcott, Trans. Amer. Ent. Soc., 48, p. 72, 1922. Indiana
British Colum-
bia, California,
Idaho, Arizona
Canada to
Florida, Utah,
California,
Arizona, Texas
Florida
California
Kansas, Utah,
Oregon, Cali-
fornia, Arizona,
New Mexico,
Texas
Arizona
Canada, British
Columbia, east
to Michigan,
south to Mexico
80 FIELDIANA: ZOOLOGY, VOLUME 32
liljebladi Wolcott, Trans. Amer. Ent. Soc., 48, p. 73, 1922. New York,
Michigan,
Wisconsin,
Pennsylvania
longissimus Wolcott, loc. cit., p. 68. New Mexico
lunatus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, New York to
p. 294, 1842. Florida, west to
bicolor Melsheimer, Proc. Acad. Nat. Sci. Phila., 2, p. 307, ISf^f8 and
1846.
mexicanus Castelnau, Silberm. Rev. Ent., 4, p. 44, 1836. Texas to
assimilis Spinola, Monog. Cler., 1, p. 249, 1844.
var. assimilis Chevrolat, Guer. Mag. Zool., (2), 5, p. 17,
1843.
var. semiochraceus Chevrolat, Rev. Mag. Zool., (3), 2,
p. 292, 1874.
var. silbermanni Chevrolat, Guer. Mag. Zool., (2), 5, p. 21,
1843.
moestus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, Oregon, South
p. 298, 1842. Dakota, Colo-
truncatus LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 23,
1 Q A Q
4y- Mexico, Ari-
zona, Texas
muttkowskii Wolcott, Bull. Wise. Nat. Hist. Soc., 7, p. 98, Michigan,
1909. Wisconsin,
Vermont
nigrifrons Say, Journ. Acad. Nat. Sci. Phila., 3, p. 190, 1823. Canada to Ala-
bama, west to
Nebraska
var. testaceipes Wolcott, Trans. Amer. Ent. Soc., 38, p. 72, Indiana
1922.
nigripes Say, Journ. Acad. Nat. Sci. Phila., 3, p. 191, 1823. Maine to Dis-
quadriguttatus auctorum (not Olivier 1795, LeConte 1849, J?ct °i5v°,!Um~
Schenkling 1906, Wolcott 1922). California
var. dubius Spinola, Monog. Cle>., 1, p. 264, pi. 25, fig. 4, south to
1844. Arizona and
var. rufiventris Spinola, loc. cit., p. 264, pi. 23, fig..3. Mexico
incerttis LeConte, Smiths. Misc. Coll., 6, No. 140, 2d ed.,
p. 55, 1866.
ocreatus Horn, Trans. Amer. Ent. Soc., 12, p. 154, 1885. Colorado to
California, New
Mexico, Ari-
zona
opifex Gorham, Biol. Centr.-Amer., Ins.-Coleop., 3, pt. 2, Texas, Mexico
p. 156, pi. 8, fig. 3, 1882.
palmii Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 218, 1904. Kansas, Utah,
New Mexico,
Arizona, Texas,
Mexico
pinus Schaeffer, Bull. Brooklyn Mus., 1, No. 7, p. 155, 1905. Arizona
WOLCOTT: FAMILY CLERIDAE
81
quadriguttatus Olivier, Ent., 4, No. 76, p. 18, pi. 2, fig. 22a, b,
1795.
quadrisignatus Say, Boston Journ. Nat. Hist., 1, p. 162,
1835.
rufescens LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 212,
1852.
var. affiliatus LeConte, Proc. Acad. Nat. Sci. Phila., 10,
p. 72, 1858.
var. latecinctus LeConte, loc. cit.
quercus Schaeffer, Bull. Brooklyn Mus., 1, No. 7, p. 154,
1905.
rosmarus Say, Journ. Acad. Nat. Sci. Phila., 3, p. 190, 1823.
angustus LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 21,
1849.
oculatus Spinola, Monog. Cler., 1, p. 267, pi. 26, fig. 1,
1844.
var. virginiensis Schaeffer, Journ. N. Y. Ent. Soc., 25,
p. 131, 1917.
rufulus Schaeffer, loc. cit.
sphegeus Fabricius, Mant. Ins., 1, p. 125, 1787.
sobrius Walker in Lord, Nat. Vane., 2, p. 326, 1866.
sobrinus Leng, Cat. Coleop. N. Amer., p. 150, 1920
(err. typ.).
var. arachnodes Klug, Abh. Konig. Akad. Wiss. Berlin,
1840-42, p. 299, 1842.
venator Chevrolat, Gue"r. Mag. Zool., (2), 5, p. 18, 1843.
lesueurii Spinola, Monog. C16r., 1, p. 249, pi. 23, fig. 2,
1844.
vetus Wolcott, Coleop. Contr., 1, p. 107, 1927.
crabronarius auctorum (not Spinola 1844).
viduus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, p. 297,
1842.
cuprescens Gorham, Cist. Ent., 2, p. 81, 1876.
erythrogaster Spinola, Monog. Cle>., 1, p. 272, pi. 27,
fig. 3, 1844.
rufiventris Chevrolat, Guer. Mag. Zool., (2), 5, p. 11,
1843 (nom. nud.).
var. jouteli Leng, Journ. N. Y. Ent. Soc., 17, p. 103, 1909.
?North America
("Caroline")
Pennsylvania
to Georgia,
west to Kansas
and Texas
Kansas, Colo-
rado, Okla-
homa, Texas,
Lower Cali- •
fornia
Arizona, Texas,
Lower Cali-
fornia
Arizona
Maine to Flor-
ida, Oklahoma,
Nebraska,
Texas to Costa
Rica
British Colum-
bia to Califor-
nia, Colorado
to Arizona and
New Mexico
Texas, Mexico
Texas
Texas, Mexico
North Carolina,
Georgia
Genus Trichodes Herbst
Trichodes Herbst, Natursyst. Ins., Kafer, 4, p. 154, 1792.
82
FIELDIANA: ZOOLOGY, VOLUME 32
Clerus Curtis, Brit. Ent., 1, pi. 44, 1824.
Genotype: Clerus alvearius Fabricius, Ent. Syst., 1, pt. 1, p. 209, 1792 (Europe)
apivorus Germar, Ins. Spec. Nov., 1, p. 81, 1824.
nutalli Say, Boston Journ. Nat. Hist., 1, p. 164, 1835
(not Kirby 1818).
var. borealis Wolcott and Chapin, Bull. Brooklyn Ent. Soc.,
13, p. 108, 1918.
interruptus LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 18,
1849 (not Klug 1842).
basalis Van Dyke, Pan-Pac. Ent., 19, p. 41, 1943.
Massachusetts
to Florida,
Alabama,
Nebraska,
Texas, Mexico
bibalteatus LeConte, Journ. Acad. Nat. Sci. Phila., (2), 4,
p. 18, 1858.
bicinctus Green, Ent. News, 28, p. 367, 1917.
bimaculatus LeConte, Trans. Amer. Ent. Soc., 5, p. 63, 1874.
bisignatus Horn, op. cit., 5, p. 231, 1876.
horni Wolcott and Chapin, Bull. Brooklyn Ent. Soc., 13,
p. 107, 1918.
illustris Horn, Trans. Amer. Ent. Soc., 5, p. 231, fig., 1876
(not Klug 1842).
nexus Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 372,
pi. 5, fig. 5, 1910.
nutalli Kirby, Trans. Linn. Soc. London, 12, p. 394, 1818.
nuthalli Spinola, Monog. Cler., 1, p. 317, pi. 31, fig. 2,
1844.
nuttalli auctorum. «
oresterus Wolcott, Field Mus. Nat. Hist., Zool. Ser., 7, p. 368,
pi. 5, fig. 4, pi. 6, fig. 21, 1910.
ornatus Say, Journ. Acad. Nat. Sci. Phila., 3, p. 189, 1823.
douglasianus White, Nomencl. Coleop. Ins. Brit. Mus., 4,
p. 60, 1849.
hartwegianus White, loc. cit.
ienellus LeConte, Proc. Acad. Nat. Sci. Phila., 10, p. 72,
1858.
var. obsoktus Wolcott, Pan-Pac. Ent., 20, p. 59, 1944.
peninsularis Horn, Proc. Calif. Acad. Sci., (2), 4, p. 382,
pi. 8, fig. 7, 1904.
simulator Horn, Trans. Amer. Ent. Soc., 8, p. 149, 1880.
flarescens Cockerell, Bull. New Mex. Exp. Sta., 28, p. 155,
1898.
California
Oklahoma,
Texas
Texas
California
Arizona, Lower
California
Lower Cali-
fornia
Maine to
Florida, Michi-
gan, Minnesota,
South Dakota,
Colorado
Texas
Alaska, British
Columbia,
Alberta, Mon-
tana, Utah,
Nevada, Ari-
zona, New
Mexico, Texas
Arizona
Arizona, Lower
California
Colorado, Utah,
Wyoming,
Arizona,
New Mexico
Genus Aulicus Spinola
Aulifus Spinola, Monog. Cle>., 1, p. 328, 1844.
Genotype: -A ulicus nero Spinola
WOLCOTT: FAMILY CLERIDAE 83
bicinctus Linsley, Univ. Calif. Publ. Ent., 6, p. 255, fig. 12, California
1936.
dentipes Schaeffer, Proc. U. S. Nat. Mus., 59, p. 157, 1921. California,
Arizona, New
Mexico, Texas
femoralis Schaeffer, Journ. N. Y. Ent. Soc., 25, p. 132, 1917. Arizona, Texas
fissipes Schaeffer, Proc. U. S. Nat. Mus., 59, p. 155, 1921. Arizona; Lower
California
humeralis Linsley, Univ. Calif. Publ. Ent., 6, p. 252, fig. 2, Lower Califor-
1936. nia, Gulf of
California,
Mexico
monticola Gorham, Biol. Centr.-Amer., Ins.-Coleop., 3, pt. 2, Arizona, Texas,
p. 146, pi. 8, fig. 18, 1882. Mexico
nero Spinola, Monog. C16r., 1, pp. 330, 331, pi. 27, fig. 5, 1844. Mexico,
?Southern or
Lower
California
nigriventris Schaeffer, Proc. U. S. Nat. Mus., 59, p. 156, 1921. Arizona,
Mexico
terrestris Linsley, Pan-Pac. Ent., 9, p. 95, 1933. California
Genus Serriger Spinola
Serriger Spinola, Monog. Cler., 1, p. 170, 1844.
Genotype: Serriger reichei Spinola
reichei Spinola, Monog. Cler., 1, p. 171, pi. 12, fig. 3, 1844. Idaho, Cali-
antennatus Schaeffer, Proc. U. S. Nat. Mus., 59, p. 158, fornia> Mexico
1921.
Genus Xenoclerus Schenkling
Xenoclerus Schenkling, Bull. Mus. Hist. Nat. Paris, 8, p. 327, 1902.
Genotype: Trogodendron edwardsi Horn
edwardsi Horn, Trans. Amer. Ent. Soc., 8, p. 149, pi. 3, fig. 8, Arizona,
1880. California
Subfamily EPIPHLOEINAE
Genus Phlogistosternus Wolcott
Phlogistosternus Wolcott, Journ. N. Y. Ent. Soc., 52, p. 124, 1944.
Phyllobaenus Spinola, Monog. Cler., 2, p. 1, 1844 (not Dejean 1837).
Genotype: Enoplium dislocatum Say
californicus Van Dyke, Bull. Brooklyn Ent. Soc., 18, p. 47, • California
1923.
84 FIELDIANA: ZOOLOGY, VOLUME 32
dislocatus Say, Journ. Acad. Nat. Sci. Phila., 5, p. 176, 1825. Canada to
distrophus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840- SSJ
.«•» „ W7A 1QjlO WMlOUli J.H1I1U1&
42, p. 374, 1842. Kansas, Colo-
transversalis Spinola, Monog. Cler., 2, p. 4, pi. 40, fig. 6, rado, Texas
1844.
merkeli Horn, Proc. Calif. Acad. Sci., (2), 6, p. 374, 1896. Arizona, New
Mexico
\
Genus Neichnea Wolcott and Chapin
Neichnea Wolcott and Chapin, Bull. Brooklyn Ent. Soc., 13, p. 108, 1918.
Ichnea Castelnau, Silberm. Rev. Ent., 4, p. 55, 1836 (part).
' Ellipotoma Wolcott in Blatchley, Bull. Ind. Dept. Geol. Nat. Res., 1, p. 858,
1910 (not Spinola 1844).
Genotype: Enoplium laticornis Say
laticornis Say, Boston Journ. Nat. Hist., 1, p. 164, 1835. Canada to
North Caro-
lina, Illinois
Genus Ichnea Castelnau
Ichnea Castelnau, Silberm. Rev. Ent., 4, p. 55, 1836.
Genotype: Ichnea lycoides Castelnau, loc. cit., p. 55 (Brazil)
elongata Knull, Ann. Ent. Soc. Amer., 32, p. 27, pi. 1, fig. 4, Arizona
1939.
Subfamily ENOPLIINAE
Genus Orthopleura Spinola
Orthopleura Spinola, Monog. Cler., 2, p. 80, 1844.
Dermestoides Schaeffer, Icon. Ins., 3, pi. 220, fig. 4, 1771.
Genotype: Tillus damicornis Fabricius
damicornis Fabricius, Suppl. Ent. Syst., p. 117, 1798. New York to
nigripennis Spinola, Monog. Cle>., 2, p. 82, 1844. nds^Kansas
tpennsylvanica Chevrolat, Rev. Mag. Zool., (3), 2, p. 325, Texas, Lower
1874. California,
thoracica Say, Journ. Acad. Nat. Sci. Phila., 3, p. 188, Mexico
1823.
punctatissima Chevrolat, Ann. Soc. Ent. France, (2), 1, p. 34, Texas
1843.
bimaculata Melsheimer, Proc. Acad. Nat. Sci. Phila., 2,
p. 307, 1846.
•
texana Bland, Proc. Ent. Soc. Phila., 1, p. 356, 1863. Texas
WOLCOTT: FAMILY CLERIDAE
85
Genus Pelonides Kuwert
Pelonides Kuwert, Ann. Soc. Ent. Belg., 38, p. 8, 1894.
Pyticara Spinola, Rev. Zool., 4, p. 75, 1841 (nom. mid.).
Genotype: Enoplium quadripunctatum Say
granulatipennis Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 220,
1904.
humeralis Horn, Trans. Amer. Ent. Soc., 2, p. 135, 1868.
militaris Chevrolat, Rev. Mag. Zool., (3), 2, p. 324, 1874.
var. perroudi Pic, L'Echange, 49, p. 12, 1933.
nigrescens Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 221, 1904.
quadripunctatus Say, Journ. Acad. Nat. Sci. Phila., 3, p. 188,
1823.
var. quadrinotatus Haldeman, Proc. Acad. Nat. Sci. Phila.,
6, p. 362, 1853.
scabripennis LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed.,
p. 98, 1866.
similis Knull, Ohio Journ. Sci., 38, p. 97, 1938.
Arizona, New
Mexico, Texas,
Mexico
New Mexico,
Texas, Mexico
Texas
Texas
Ohio, Indiana,
Arkansas,
Kansas, Texas
Lower Cali-
fornia
Texas
Genus Chariessa Perty
Chariessa Perty, Del. Anim. Artie., p. 109, 1830.
Brachymorphus Chevrolat, Coleop. Mexico, Cent. II, Fasc. 7, No. 150, 1835.
Pelonium Spinola, Monog. Cler., 1, p. 347, 1844 (part).
Philyra Castelnau, Silberm. Rev. Ent., 4, p. 53, 1836.
Tarandocerus Chevrolat, Mem. Cle>., p. 7, 1876.
Genotype: Chariessa ramicornis Perty, Del. Anim. Artie., p.
1830 (Brazil)
dichroa LeConte, Repts. Expl. Surv. Miss.-Pacif., 12, Zool.
Rept. No. 1, p. 48, 1859.
elegans Horn, Trans. Amer. Ent. Soc., 3, p. 87, 1870.
lemberti Rivers, Zoe, 6, p. 396, 1894.
floridana Schaeffer, Journ. N. Y. Ent. Soc., 25, p. 133, 1917.
pilosa Forster, Nov. Spec. Ins., p. 49, 1781.
var. marginata Say, Journ. Acad. Nat. Sci. Phila., 3, p. 187,
1823.
cincta Spinola, Monog. Cler., 1, p. 358, pi. 36, fig. 4, 1844.
onusta Say, Amer. Ent., 3, pi. 41, fig. 1, 1828.
texana Wolcott, Ent. News, 19, pp. 70 (key), 72, 1908.
109, pi. 22, fig. 3,
California
British Colum-
bia, Oregon,
California,
Texas
Florida (Key
Largo)
Canada to
Florida, west to
Montana,
Kansas, Okla-
homa, and
Texas
Texas
86
FIELDIANA: ZOOLOGY, VOLUME 32
vestita Chevrolat, Coleop. Mexico, Cent. II, Fasc. 7, No. 150, Texas, Mexico,
1835. Costa Rica
spectabilis Castelnau, Silberm. Rev. Ent., 4, p. 50, 1836.
Genus Corinthiscus Fairmaire and Germain
Corinthiscus Fairmaire and Germain, Coleop. Chilensia, II, p. 4, 1861.
Pelonium Spinola, Monog. Cler., 1, p. 347, 1844 (part).
Genotype: Corinthiecus insignicornis Fairmaire and Germain, Coleop. Chilen-
sia, II, p. 4, 1861 (Chile)
fasciatus LeConte, Ann. Lye. Nat. Hist. N. Y., 5, p. 214, 1852. California,
Texas
var. trilobatus Van Dyke, Bull. Brooklyn Ent. Soc., 10, California
p. 32, 1915.
granulosus Wolcott, Bull. Wise. Nat. Hist. Soc., 7, p. 100, Texas
1909.
leucophaeus Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, District of
p. 366, 1842. Columbia to
vetustus Spinola, Monog. Cler., 1, p. 360, pi. 35, fig. 4,
souri, Kansas,
albomaculatus Ziegler, Proc. Acad. Nat. Sci. Phila., 2, Texas, Lower
p. 268, 1845. California
venustus Haldeman, op. cit., 3, p. 126, 1847.
maculicollis Schaeffer, Journ. N. Y. Ent. Soc., 12, p. 219, Texas
1904.
peninsularis Schaeffer, op. cit., 25, p. 132, 1917.
Lower Cali-
fornia
Genus Cregya LeConte
Cregya LeConte, Smiths. Misc. Coll., 3, No. 136, 1st ed., p. 197, 1861.
Galeruclerus Gahan, Ann. Mag. Nat. Hist., (8), 5, p. 75, 1910.
Genotype: Clerus oculatus Say
mixta LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed., p. 98, Maryland to
1866. Louisiana,
Ohio, Ken-
tucky, Texas
oculata Say, Boston Journ. Nat. Hist., 1, p. 163, 1835.
rginipennis
fig. 6, 1844.
Massachusetts,
marginipennis Spinola, Monog. Cler., 1, p. 363, pi. 35, District of
ficr « i»/i4 v/onimraa,
Louisiana,
Illinois, Kansas,
Texas
quadrisignata Spinola, loc. cit., p. 365, pi. 33, fig. 3.
Texas, Central
America
var. bilineicollis Chevrolat, Rev. Mag. Zool., (3), 2, p. 327, Texas
1874.
WOLCOTT: FAMILY CLERIDAE 87
quadrinotata Chevrolat, loc. cit., p. 328.
uncinata Chevrolat, loc. cit.
Subfamily KORYNETINAE1
Genus Tarsostenus Spinola
Tarsostenus Spinola, Monog. C16r., 1, p. 287, 1844.
Genotype: Clerus univittatus Rossi
univittatus Rossi, Fauna Etrusc., 1, p. 44, 1792. Cosmopolitan
albofasciatus Melsheimer, Proc. Acad. Nat. Sci. Phila., 2,
p. 306, 1846.
biguttatus Montrouzier, Ann. Soc. Ent. France, (3), 8,
p. 260, 1860.
fasdatellus Spinola, Monog. Cle>., 2, p. 172, pi. 8, fig. 5,
1844.
fasciatus Curtis, Brit. Ent., 6, pi. 270, 1832.
moerens Westwood in White, Nomencl. Coleop. Ins. Brit.
Mus., 4, p. 57, 1849.
picipennis Westwood in White, loc. cit., p. 48.
succinctiis Chevrolat, Rev. Zool., 5, p. 277, 1842.
Genus Lebasiella Spinola
Lebasiella Spinola, Monog. Cle"r., 2, p. 77, 1844.
Genotype: Lebasiella erythrodera Spinola,2 .loc. cit., p. 79, pi. 43, fig. 1 (Colom-
bia)
discolor Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, Texas, Mexico
p. 353, 1842.
var. tibialis Schenkling, Deutsch. Ent. Zeitschr., 1898, California,
p. 368, 1898. Mexico
marginella Chevrolat, Ann. Soc. Ent. France, (2), 1, p. 42, California,
1843. Mexico
pallipes Klug, Abh. Konig. Akad. Wiss. Berlin, 1840-42, Pennsylvania,
p. 353, 1842. Texas, Mexico
nigripennis LeConte, Smiths. Misc. Coll-., 6, No. 167, 2d
ed., p. 99, 1866.
lErolestus cleroides Wolcott (1927, p. 108) was described as a new genus and
species from Tioga County, Pennsylvania, on the basis of an erroneous locality
label and should not be included in the North American lists. It is a synonym
of the Australian Blackburniella intricata Blackburn.
The North American species formerly included in Korynetes are here trans-
ferred to Loedelia. Korynetes is not represented in our fauna. The only New
World species left in the genus are Chilean and further study may show that these
are improperly placed.
2 Gorham, in Trans. Ent. Soc. London, 1878, p. 156, cites Orthopleura lepida
Klug as type of Lebasiella. As lepida Klug was not originally included in the genus,
the citation is invalid.
88
FIELDIANA: ZOOLOGY, VOLUME 32
Genus Loedelia Lucas
Loedelia Lucas, Arch. Naturg., 84, Abt. A, p. 380, 1920.
Necrobioides Gahan, Ann. Mag. Nat. Hist., (8), 5, p. 76, 1910 (not Fairmaire
1882).
Genotype: Necrobioides mexicanus Gahan, loc. cit. (Mexico)
discoidea LeConte, Trans. Kans. Acad. Sci., 7, p. 77, 1881. Arizona
janthina LeConte, Smiths. Misc. Coll., 6, No. 167, 2d ed.,
p. 99, 1866.
maculicollis LeConte, Trans. Amer. Ent. Soc., 5, p. 63, 1874.
filiolus Chevrolat, Rev. Mag. Zool., (3), 2, p. 326, 1874.
lineatocolle Chevrolat, loc. cit.
var. nigricollis Wolcott, Coleop. Contr., 1, p. 110, 1927.
Lower Cali-
fornia
California
California
Genus Necrobia Olivier
Necrobia Olivier, Ent., 4, No. 76bis, p. 5, 1795.
Agonolia Mulsant and Rey, Col. France, 14, Angusticolles, p. 122, 1863.
Genotype: Dermestes violaceus Linnaeus
Pmesosternalis Schaeffer, Journ. N. Y. Ent. Soc., 16, p. 135,
1908.
ruficollis Fabricius, Syst. Ent., p. 57, 1775.
var. knochei Kolbe, Breslau Jahresh. Ver. Insektenk., 5, p. 9,
1912.
rufipes DeGeer, Mem., 5, p. 165, pi. 15, fig. 4, 1775.
amethystina Stephens, 111. Brit. Ent., Mandib., 5, p. 417,
1832.
dermestoides Filler and Mitterpacher, Iter Poseg., p. 68,
pi. 7, fig. 8, 1783.
foveicollis Schenkling, Mitt. Mus. Naturhist. Hamburg,
17, p. 20, 1900.
glabra Champollion, Mag. Encycl. Millin., 3, p. 41, 1814.
mumiarum Hope in Pettigrew, Hist. Egypt. Mum.,
p. 54, pi. 5, figs. 1-3, 1834.
pilifera Reitter, Verh. Nat. Ver. Briinn, 32, p. 85, 1894.
var. aeneipennis Csiki, Rovart. Lapok, 7, p. 124, 1900.
violacea Linnaeus, Syst. Nat., ed. 10, 1, p. 356, 1758.
angustata Faldermann, Nouv. M6m. Soc. Nat. Moscou, 4,
p. 210, 1835.
chalybea Sturm, Deutschl. Fauna, Ins.-Kafer, 11, p. 43,
pi. 232, figs. A-O, 1837.
errans Melsheimer, Proc. Acad. Nat. Sci. Phila., 2, p. 307,
1846.
quadra Marsham, Ent. Brit., 1, p. 323, 1802.
Arizona,
Mexico
Cosmopolitan
Cosmopolitan
Cosmopolitan
WOLCOTT: FAMILY CLERIDAE 89
Genus Opetiopalpus Spinola1
Opetiopalpus Spinola, Monog. Cler.,*2, p. 110, 1844.
Opetiopselaphus Gemminger and Harold, Cat. Coleop., 6, p. 1759, 1869.
Genotype: Clerus scutellaris Panzer
luridus Spinola, Monog. Cler., 2, p. 113, pi. 45, fig. 5, 1844. ?NorthAmerica
1 Opetiopalpus luridus Spinola is retained in the catalogue with reservations.
It seems highly probable that the species was described as being from the United
States on the basis of mislabeled specimens. Apparently neither specimens
referable to this species nor species referable to this Old World genus have been
found in the New World during the past century.
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94 FIELDIANA: ZOOLOGY, VOLUME 32
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WOLCOTT: FAMILY CLERIDAE 95
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INDEX
Current names in roman type, synonyms in italic type.
Ababa Casey, 76
abdominalis Chevr. (Enoch), 78
abdominalis Kirby (Than.), 78
abruptus LeC., 79
acerbus Wolc., 78
aegra Newm. (Isohyd.), 74
aegra Wolc. (Gym.), 68
aemula Wolc., 68
aeneicollis Spin., 79
aeneipennis Csiki, 88
aequinoctialis Spin., 74
aethiops Wolc., 68
affiliatus LeC. (Enoch), 81
affiliatus Fall (Phyll.), 71
Agonolia Muls. & Rey, 88
albocincta Horn, 74
albofasciatus Melsh., 87
albomaculatus Zieg., 86
alvearius Fabr., 82
amethystina Steph., 88
analis LeC., 78
angustata Spin. (Cym.), 68
angustata LeC. (Cym.), 68
angustata Fald. (Necr.), 88
angustus LeC., 81
antennata Schffr. (Cym.), 68
antennatus Schffr. (Serr.), 83
aprivorus Germ., 82
arachnodes Klug, 81
arizonica Schffr. (Cym.), 71
arizonicus Schffr. (Phyll.), 71
assimilis Chevr. (Enoch), 80
assimilis Spin. (Enoch), 80
Aulicus Spin., 82
badeni Gorh., 78
balteata LeC., 68
basalis Van Dyke, 82
belfragei Horn, 68
bibalteatus LeC., 82
bicinctus Green (Trich.), 82
bicinctus Lins. (Auh), 83
bicolor LeC. (Phylh), 71
bicolor Say (Cym.), 68
bicolor Melsh. (Enoch), 80
bicornus Van Dyke, 71
biguttatus Montr., 87
bilineicollis Chevr., 86
bimaculata Melsh. (Orth.), 84
bimaculatus LeC. (Trich.), 82
bimaculatus Skinner (Enoch), 78
bimaculatus Wolc. (Phylh), 72
binotatus Chpn., 72.
bioculatus Schffr., 78
bisignatus Horn, 82
Blackburniella Chpn., 87
blanchardi Wolc., 72
bombycinus Chevr., 79
borealis Wolc. & Chpn., 82
bosci Chevr., 71
Bostrichoclerus Van Dyke, 71
Brachymorphus Chevr., 85
brachyptera Klug, 74
brevicollis Schffr. (Cym.), 69
brevicollis Spin. (Enoch), 79
brunnea Chpn. (Isohyd.), 74
brunnea Melsh. jXiecont.), 68
brunnea Spin. (Cym.), 71
buquet Lef. (Than.), 75
buqueti Westw. (Than.), 75
calif ornica Fall (Mon.), 67
calif ornica Horn (Cym.), 69
calif ornica Horn (Cym.), 69
calif ornicus Van Dyke (Phlog.), 83
Callotillus Wolc., 67
cancellata LeC., 68
carbonarius Spin., 68
castanea Newm., 76
catalina Cazier, 76
cephalica Schffr., 69
chalybea Sturm, 88
chapini Wolc. (Enoch), 79
chapini Wolc. (Phylh), 72
Chariessa Perty, 85
chiricahuae Knull, 76
cinctus Spin. (Phylh), 72
cincta Spin. (Char.), 85
CLERINAE, 76
Cleroides Schffr., 77
cleroides Wolc., 87
Cleronomus Klug, 76
Clerus Curtis, 82
Clerus auct., 78
cobaltinus Chpn., 72
coccineus Schklg., 79
coeruleipennis Wolc., 72
cognata Wolc., 69
colbaltina Chpn., 72
Colyphus Spin., 76
collaris Spin., 68
comans Wolc., 69
101
102
FIELDIANA: ZOOLOGY, VOLUME 32
commixtus Chpn., 72
confusa Wolc., 69
corallinus Fall, 79
cordifer LeC., 79
Corinthiscus Fairm. & Germ., 86
crabronarius Spin. (Enoch), 79
crabronarius auct. (Enoch), 81
Cregya LeC., 86
cribripennis Fall, 72
crinita Casey, 76
cuneiformis Wolc., 72
cupressi Van Dyke, 79
cuprescens Gorh., 81
curtipennis Newm., 74
cyanescens LeC., 72
cyanipennis Klug, 77
cyanitinctus Fall, 72
Cylinder Voet, 67
cylindricollis Chevr., 69
Cymatodera Gray, 68
Cymatoderus Desmarest, 68
damicornis Spin., 84
dasytoides Westw., 77
decipiens Fall, 69
decussatus Klug, 79
delicatula Fall, 69
dentipes Schffr., 83
Derestenus Chevr., 77
Dermestoides Schffr., 84
dermestoides Klug (Than.), 75
dermestoides Pill. & Mitt. (Near.), 88
dichroa LeC., 85
difficilis LeC., 72
discoidea LeC. (Loed.), 88
discoideus LeC. (Phylh), 72
discolor Klug, 87
dislocatus Say, 84
distrophus Klug, 84
douglasianus White, 82
dubius Fab. (Than.), 77
dubius Wolc. (Phylh), 72
dubius Spin. (Enoch), 80
eburneocinctus Wolc., 67
edwardsi Horn, 83
Elasmocerus LeC., 67
elegans Er. (Call.), 67
elegans Horn (Char.), 85
Ellipotoma Wolc., 84
elongata Linn. (Till.), 67
elongata Knull (Ichn.), 84
Enoclerus Gahan, 78
ENOPLIINAE, 84
EPIPHLOEINAE, 83
Erolestus Wolc., 87
errans Melsh., 88
erro Wolc., 79
erythrodera Spin., 87
erythrogaster Spin., 81
eximius Mannerh., 79
fallax Wolc. (Phylh), 72
fallax Horn (Cym.), 68
fasciatellus Spin., 87
fasciatus Curtis (Tars.), 87
fasciatus LeC. (Corinth.), 86
fascifera LeC., 69
femoralis Schffr., 83
filiolus Chevr., 88
fissipes Schffr., 83
flavescens Cklh, 82
flavosignata Schffr., 69
floridana Schffr., 85
formicarius Linn., 77
foveicollis Schklg., 88
fraterna Wolc., 73
fuchsii Schffr. (Cym.), 69
fuchsi Schffr. (Phylh), 72
funebris Chevr., 73
furcata Gorh. (Phylh), 72
furcatus Schffr. (Col.), 77
fuscula LeC., 69
gahani Wolc., 72
Galeruclerus Gorh., 86
gerhardi Wolc. (Enoch), 79
gerhardi Wolc. (Isohyd.), 74
gigantea Horn, 69
girodi Chevr., 75
glabra Champ., 88
gnara Wolc., 68
granulatipennis Schffr., 85
granulosus Wolc., 86
haagi Chevr., 77
hamatus LeC., 72
hartwegianus White, 82
holosericeus White, 79
hopei Gray, 69
hopfneri Spin., 79
horni Wolc. (Cym.), 69
horni Wolc. & Chpn. (Trich.), 82
humeralis Horn (Pel.), 85
humeralis Linsley (Auh), 83
humeralis Say (Phylh), 72
humeralis Schffr. (Enoch), 79
Hydnocera Newm., 71
Ichnea Cast., 84
Ichnea Cast., 84
ichneumoneus Fab., 79
illustris Horn, 82
incertus LeC., 80
inornata Say, 69
insignicornis Fairm. & Germ., 86
interruptus LeC., 82
intricata Blackb., 87
inyoensis Van Dyke, 79
iowensis Chpn., 72
isabellae Wolc., 69
Isohydnocera Chpn., 74
janthina LeC., 88
jouteli Leng, 81
WOLCOTT: FAMILY CLERIDAE
103
knabi Wolc., 79
knausi Wickh. (Phyll.), 72
knausi Wolc. (Gym.), 69
knochei Kolbe, 88
Korynetes Herbst, 87
KORYNETINAE, 87
laetus Klug, 79
laevicollis Schffr., 69
latecinctiis LeC., 81
latefascia Schffr., 69
lateralis Schffr., 72
laticornis Say, 84
lautus Wolc., 79
Lebasiella Spin., 87
lecontei Wolc. (Enoch), 79
lecontei Wolc. (Phyll.), 72
lecontei Wolc. (Prioc.), 76
Lecontella Wolc. & Chpn., 68
lemberti Rivers, 85
lepida Klug, 87
lesueurii Spin., 81
leucophaeus Klug, 86
liebecki Wolc., 74
liljebladi Wolc., 80
limbata Spin., 74
lineatocolle Chevr. (Loed.), 88
lineatocollis Spin. (Phyll.), 74
Loedelia Lucas, 87
longicollis Spin. (Gym.), 71
longicollis Zieg. (Isohyd.), 74
longicornis LeC., 69
longissimus Wolc., 80
longula Wolc., 70
longus LeC., 72
lunatus Klug, 80
luridus Spin., 88
lycoides Cast., 84
Macrotelus Klug, 67
maculata Zieg., 76
maculicollis LeC. (Loed.), 88
maculicollis Schffr. (Corinth.), 86
marginata Gorh. (Phyll.), 74
marginata Say (Char.), 85
marginella Chevr., 87
marginipennis Spin., 86
maritimus Wolc., 73
megatoma Spin., 67
melanocephalus Chevr., 78
melanopterus Dury, 77
merkeli Horn, 84
mesosternalis Schffr., 88
Metademius Mats., 75
mexicanus Cast., 80
mexicanus Gahan, 88
militaris Chevr., 85
mima Wolc., 74
mirus Wolc., 73
mitchelli Chpn., 70
mitis Wolc., 70
mixta LeC., 86
moerens Westw., 87
moestus Klug, 80
months Melsh., 77
Monophylla Spin., 67
monticola Gorh. (Aul.), 83
monticola Wolc. (Than.), 78
morosa LeC., 70
mumiarum Hope, 88
muttkowskii Wolc., 80
mystica Wolc., 70
Necrobia Oliv., 88
Necrobioides Gahan, 88
Neichnea Wolc. & Chpn., 84
neomexicana Knull, 70
nero Spin., 83
nexus Wolc., 82
nigrescens Schffr. (Pel.), 85
nigrescens Schffr. (Phyll.), 73
nigricollis Wolc., 88
nigrifrons Say, 80
nigrina Schffr., 74
mgripennis LeC. (Leb.), 87
nigripennis Spin. (Orth.), 84
nigripes Say (Enoch), 80
nigripes var. White (Than.), 78
nigriventris Schffr. (Aul.), 83
nigriventris LeC. (Enoch), 79
niveifasciatus Schffr., 73
nubilus Klug, 78
nunnenmacheri Schffr., 72
nutalli Kirby, 82
nutalli Say, 82
nuttalli auct., 82
nutthalli Spin., 82
obliquefasciata Schffr., 70
oblita Horn, 70
pbsoletus Wolc., 82
occidentals Chpn. (Phyll. ), 73
occidentalis Gorh. (Call.), 67
ocreatus Horn, 80
oculata Say (Greg.), 86
oculatus Spin. (Enoch), 81
omoger Horn, 73
onusta Say, 85
Opetiopalpus Spin., 89
Opetiopselaphus Gemm. & Har., 89
opifex Gorh., 80
oresterus Wolc., 82
ornata Wolc. (Isohyd.), 74
ornaticollis LeC., 77
ornatus Say (Trich.), 82
Orthopleura Spin., 84
ovipennis LeC., 70
pallida Schffr., 70
pallipennis Say, 73
pallipes Klug (Leb.), 87
pallipes Schffr. (Mon.), 67
pallipes Wolc. (Plac.), 77
palmii Schffr., 80
104
FIELDIANA: ZOOLOGY, VOLUME 32
parviceps Schffr., 75
patagoniae Knull, 68
pedalis LeC., 75
Pelonides Kuw., 85
Pelonium Spin., 85, 86
peninsularis Horn (Trich.), 82
peninsularis Schffr. (Corinth.), 86
peninsularis Schffr. (Cym.), 70
pennsylvanica Chevr., 84
Perilypus Spin., 68
perroudi Pic, 85
Philyra Cast., 85
Phloeopterus Britt., 77
Phlogistosternus Wolc., 83
PHYLLOBAENINAE, 71
Phyllobaenus Dej., 71
Phyllobaenus Spin., 83
picipennis Chpn. (Phyll.), 73
picipennis Westw. (Tars.), 87
pilifera Reitt., 88
pilosa Forster, 85
pilosella LeC., 70
pinus Schffr., 80
plagifer Fall, 73
Placopterus Wolc., 77
Ploeopterus Leng, 77
plumbea Gorh., 77
Poecilochroa Chevr., 77
Pripcera Kirby, 76
Prionodera Wolc., 76
Prionostichaeus Wolc., 76
Pseudoclerus J. du Val, 77
pubescens LeC. (Phyll.), 73
pubescens Wolc. (Cym.), 70
pulcher Chpn., 73
punctata LeC. (Cym.), 70
punctata Spin. (Phyll.), 74
punctatissima Chevr., 84
puncticollis Bland, 70
puritanus Wolc., 73
purpuricollis Horn, 70
pusilla Kirby (Prioc.), 76
pusilla Schffr. (Isohyd.), 74
pygmaeus Wolc., 73
Pyticara Spin., 85
quadra Marsham, 88
quadriguttatus Oliv. (Enoch), 81
quadriguttatus auct. (Enoch), 80
quadrimaculatus Van Dyke, 73
quadrinotala Chevr. (Greg.), 87
quadrinotatus Hald. (Pel.), 85
quadripunctatus Say, 85
quadrisignata Spin. (Creg.), 86
quadrisignatus Say (Enoch), 81
quercus Schffr., 81
ramicornis Perty, 85
reichei Spin., 83
repandus Horn, 78
robustus Horn, 73
rosmarus Say, 81
rubriventris LeC., 78
rudis Wolc., 70
rufescens LeC. (Enoch), 81
rufescens Spin. (Prioc.), 76
ruficeps Spin., 77
ruficollis Fahr. (Necr.), 88
ruficollis Schffr. (Mon.), 67
rufipes DeGeer (Necr.), 88
rufipes Newm. (Phylh), 73
rufiventris Wolc. (Cym.), 70
rufiventris Chevr. (Enoch), 81
rufiventris Spin. (Enoch), 80
rufulus Schffr., 81
rufus Oliv., 79
sanguineus Say, 76
santarosae Schffr., 70
scaber LeC., 73
scabripennis LeC., 85
schusteri LeC., 74
schwarzi Wolc., 70
scutellaris Panzer, 89
semiochraceus Chevr., 80
sericans Westw., 79
serrata Newm., 73
Serriger Spin., 83
sexguttatus Fabr., 78
signaticollis Spin., 77
stlbermanni Chevr., 80
similis Knull, 85
simulans Schffr., 73
simulator Horn, 82
singularis Wolc., 73
sirpata Horn, 70
snowi Wolc., 70
sobrina Fall (Wolc.), 75
sobrinus Leng (Enoch), 81
sobrius Walker, 81
soror Wolc., 70
spatiosa Wolc., 70
spectabilis Cast., 86
sphegeus Fabr., 81
spinolae LeC., 78
steniformis Spin., 73
subaena LeC. (Phylh), 72
subaenus Spin. (Phylh), 73
subcostatus Schffr., 77
subfasciatus LeC., 73
subsimilis Wolc., 70
substriata Wolc., 67
succinctus Chevr., 87
suturalis Klug, 73
tabacci Mats., 75
tabida LeC., 75
Taneroclerus Chevr., 75
tantilla LeC., 76
Tarandocerus Chevr., 85
Tarsostenus Spin., 87
tenella Say (Phylh), 74
tenellus LeC. (Trich.), 82
tenera LeC., 69
WOLCOTT: FAMILY CLERIDAE
105
terminata Say, 67
terrestris Linsley, 83
testaceipes Wolc., 80
texana Bland (Orth.), 84
texana Wolc. (Char.), 85
texana Gorh. (Cym.), 69
Thanasimus Latr., 77
Thanateroclerus Gemm. & Har., 75
Thaneclerus Desmarest, 75
THANEROCLERINAE, 75
Thaneroclerus Lef., 75
Thea.no Chevr., 71
thoracica Say (Orth.), 84
thoracicus Oliv. (Plac.), 77
tibialis Wolc. (Phyll.), 74
tibialis Schklg. (Leb.), 87
TILLINAE, 67
Tillus Oliv., 67
torosa Wolc., 71
transversalis Spin., 84
Trichodes Hbst., 81
tricolor Schffr. (Phyll.), 74
tricolor Skin. (Cym.), 71
tricondylae LeC., 75
trifasciatus Say, 78
trilobatus Van Dyke, 86
tristis Schffr., 74
truncatus LeC., 80
turbata Horn, 71
tuta Wolc., 71
umbrina Fall, 71
uncinata Chevr., 87
undatulus Say, 78
undulata Say (Cym.), 71
undulatus Leng et auct. (Than.), 78
unifasciatus Say, 74
uniformis Schffr., 71
univittatus Rossi, 87
usta LeC., 71
vafer Wolc., 67
van dykei Schffr., 71
variegata Kirby, 76
vauthieri Spin., 71
venator Chevr., 81
venustus Hald., 86
verticalis Say, 74
vestita Chevr., 86
vetus Wolc., 81
vetiistus Spin., 86
vicinus Chpn., 74
viduus Klug, 81
violacea Linn., 88
virginiensis Schffr., 81
wickhami Wolc., 74
Wolcottella Lucas, 76
Wolcottia Chpn., 75
xanti Horn, 71
xavierae Knull, 71
Xenoclerus Schklg., 83
Xenodosus Champl., 76
Zenodosus Wolc., 75
Zenodusus Leng, 76
zonatus Klug, 78
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